Life itself
Evolution's requisite forces & mechanisms are typically described as 1) selective pressure upon populations 2) discrete genetic inheritance and 3) genetic mutation and recombination.
This list falls far short, and will be looked at as inadequate, by evolutionary biologists in the late 21st century.
Number 1 on the list should probably be life itself: a reliable ontogeny, or morphological unfolding, based upon differentiating steps under the guidance of genetic information. This would not take place without the power of a stunningly robust, coherent developmental physiology, which essentially always creates a functioning life.
Life's doggedness is quite tangible, yet the nature of its force & robustness is still a mystery. The issue is so overarching as to be barely ever mentioned, except by specialists working on the origins of life, and then only occasionally. Without this force of life, natural selection could not take place, genetic information would have nothing to guide, and genetic mutation & recombination would be moot.
I believe there's some scientific cultural artifact that makes discussion of a life force somehow sound religious. I'm not religious, but I'm given funny looks when I say "life force". One of the most basic biological facts is regularly ignored by the scientific establishment, much to the impoverishment of scientific discussion. I suppose since it is not patentable, it is ignored in these times of fame & fortune. It's the elephant at the dinner table.
